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He has two sons,and ( ) are collage students.

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He has two sons,and ( ) are collage students.
为什么能用both of them,and前后可以用两个主语么
如果是
he has two sons,___are college students.
逗号前后只能有一个主语,
能用both of them,and前后可以用两个主语.
如果是
he has two sons,both of whom are college students.